The painter James Whistler used dark colors to emphasize the melancholy mood of his works, best known as a tonalist, and played on haunting shadows in his most well-known piece, "Whistler's Mother."
A) The painter James Whistler used dark colors to emphasize the melancholy mood of his works, best known as a tonalist, and
B) The painter James Whistler used dark colors to emphasize the melancholy mood of his works, being a tonalist, and
C) The painter James Whistler, best known as a tonalist, used dark colors to emphasize the melancholy mood of his works, and
D) The painter James Whistler, best known as a tonalist, used dark colors to emphasize the melancholy mood of his works, which he
E) The painter James Whistler, best known as a tonalist, used to emphasize the melancholy mood of his works with dark colors, and
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